Mesic to dry wood and grassy meadows in the lowland and montane zones (Douglas et al., 1999); Moist, often saline meadows, wet coastal rocks and bluffs, stream banks (Pojar and Mackinnon, 1994); Sporadic in open-canopy forests on water-shedding sites, or in meadow-like communities where early spring moisture is followed by mid-summer drought. Occasionally inhabits water-receiving sites (vernal springs) (Klinka et al., 1989).
Nitrogen-rich soils (Klinka et al., 1989).
Dry to moderately moist soils (Klinka et al., 1989).
Shade tolerant/intolerant (Klinka et al., 1989).