Moist (at least in spring), open to shady, often sandy, forests, thickets, meadows. Common at low to medium elevations (Pojar and MacKinnon, 1994). Southern B.C. east to Alberta and south through the Rocky Mountain and Pacific states to California (Young, 2001).
Grows on a variety of substrates including sand, riverine silts, gravel, rock crevices, talus and scree; also found on burned sites (USDA, Forest Service Fire Effects Information System).
Dry to fresh (Pojar and MacKinnon, 1994).
Shade tolerant on drier sites, moderately shade intolerant on moist sites (USDA, Forest Service Fire Effects Information System).
Facultative seral species. Occurs at all seral stages, often colonizing disturbed sites after fire (USDA, Forest Service Fire Effects Information System).
